Professor Yossi Loya

State University of New York at Stony Brook, L.I. New York, 1967-1971.


(Office) +972-3-640-7683
(Fax) +972-3-640-7682

(Home) +972-9-774-1921
Mobile: 054-637-2121

Room#: 326
Member's portrait
  Personal Information
  Research Interests
  Selected Publications
  Students and Lab Members
  Academic and Professional Experience
  Full CV
  My Theme

Personal Information
Home Adress: 31 Hakochav St. Raanana, 43568 Israel
    Post   doctorate                
Woods Hole Oceanographic Institution, (WHOI) Mass.

Oil pollution effects on benthic communities

in Buzzards Bay Woods Hole



State University of New York at Stony Brook, New York


Community structure and species diversity

of hermatypic corals at Eilat, Red Sea.

Supervisor: Prof. L.B. Slobodkin



Tel Aviv University,



Ecology of fish breeding in brackish water ponds near the Dead Sea (Israel); Suma cum laude

Tel Aviv University, Israel

Research Interests

Ecology and Evolution of reef corals; Coral reef community structure; Species diversity and community structure of corals; Life history strategies of reef corals and their associated fauna; Effects of environmental stress on anthozoan reproduction; Sex change in corals; Hormonal influences on anthozoan reproduction Competitive networks and space partitioning within coral populations; The role of boring organisms in bioerosion of coral reefs; Effects of oil-pollution on coral- reef communities; Effects of natural and anthropogenic disturbances on coral reef communities; Conservation ecology and management of coral reef nature reserves; Biologically active compounds derived from coral-reef organisms in search for new drugs; Global climatic changes and its effect on coral-reef communities; Coral bleaching; Coral diseases.

Students and Lab Members


                                                          Ph.D. students

Graduation year




                       Thesis title

Yehuda Benayahu

Population dynamics and life history strategies in Red Sea soft corals.

Baruch Rinkevich

Calcification and productivity in the scleractinian coral Stylophora pistillata.

Avi Shafir

Aspects of energy flow within the coral Stylophora pistillata and some of

its associates.

Yechiam Schlesinger
Reproduction and juvenile growth in stony corals.
Craig Browdy

Growth and reproduction of the shrimp Penaeus semisulcatus in captivity.

Micha Ilan

 Life strategies of sponges from the Red Sea: Reproduction, settlement and

self/non-self recognition.

Avigdor Abelson

Biomechanical aspects in the biology and ecology of sessile organisms in coral reefs.

Ofer Mokady

Bioerosion rate of corals by boring bivalves: A chemical approach.

Rami Klein
Skeletal banding in recent and fossil corals.
Amit Lotan
Medusae blooms in the Mediterranean Sea: Ecological and toxicological aspects.
Ester Kramarsky
Winter Reproductive strategies of fungid corals: sexual and asexual reproduction.
Uri Oren
Regeneration patterns in reef corals.
Ariel Kushmaro

Interactions between corals and their associated  bacteria

(Co-Supervisor: Prof. Eugene Rosenberg).

Maoz Fine

Community structure and dynamics of Mediterranean corals.

Michael Rosenfeld

The use of contemporary corals in predictive models concerned with

global climate change.

Dov Kelman

Biologically active materials derived from Red Sea organisms.

Omer Choresh

Expression of heat shock proteins (HSPs) in marine invertebrates:development

of an early warning system for disturbed marine environments.

Gidon Winters

Photoinhibition in corals – effects of UV, PAR and temperature.

Noa Shenkar

Population dynamics of Mediterranean and Red Sea tunicates.

Amy Shlesinger

Nudibranch / Anemone associations: a study of life histories and interactions.

Assaf Zvuloni

Spatial patterns in coral reef populations.

Rachel Armosa

Physiological mechanisms involved in the reproduction of Anthozoa.

Omri Bronstein

Ecology, biology, and taxonomy of coral-reef associated
sea urchins in the Red Sea and western Indian Ocean

Ada Almaru

Molecular systematics of benthic ctenophores (Family: Coeloplanidae)

from the Gulf of Aqaba, Red Sea.

Gal Eyal
Biodiversity of Mesophotic Scleractinian coral in the Gulf of Eilat/Aqaba.
Lee Eyal Shacham   Reproductive effort of mesophotic corals (60 m depth) and shallow reef corals.
Tom Schlesinger Connectivity between mesophotic corals  (60 m depth) and shallow reef corals in the Gulf of Eilat/Aqaba.


                                               M.Sc. students

Graduation year




                               Thesis title

Yehuda Benayahu

Quantitative characteristics of stony corals, soft corals and algae in the northern Gulf of Eilat ( Red Sea ).

Baruch Rinkevich

On the reproduction of Stylophora pistillata (Esper) and harmful effects of oil pollution on its population.


Yechiam Schlesinger

Effects of phosphate pollution on the community structure of the holothurians Actinopyga bannwarthi and Synapta maculata.

Mordehai Shpigel

Spatial heterogeneity in branching corals and fish species diversity.


Zeev Wolodarsky

Competition and space partitioning among Trapezia species within the coral Stylophora pistillata .


Esther Kramarsky -


Life history strategy of the bivalve Lithophaga purpurea boring in the coral Cyphastrea chalcidicum.

Itzchak Brikner

Reproductive and settlement strategy of the boring bivalve Lithophaga purpurea in living corals.

Avigdor Abelson

Aggressiveness in stony corals: Is it competition for space?

Oved Gur

Predation and life history strategies of the coral predatory snail Drupella cornus.

Sara Sadot Life history and reproduction of Pocillopora in the Gulf of Eilat.  
Rami Klein

Sclerochrondogical aspects of hermatypic corals (genus: Porites) in the Gulf  of Eilat.

Gila Arazi
Larval ecology of boring bivalves in living corals.
Oron Prager

Surface structural complexity and its influence on ecological indices of aquatic communities.


Tamar Liberman


Possible benefits to the coral Stylophora pistillata from the association with the fish Dascyllus marginatus

Nadav Shashar
Nitrogen fixation in stony corals.
Vered Shimoni
Population genetics of boring bivalves in stony corals.
Dalit Trovezky

Bioerosion of the coral reef by parrotfish

(Co-supervisor: B. Lazar , Hebrew University ).

Maoz Fine

The scleractinian coral Oculina patagonica : A new invader to the Mediterranean sea, biological and ecological aspects.

Hadas Lubinevsky
Light and shade adapted Mediterranean corals
Omer Choresh
Heat shock proteins in Oculina patagonica: a Mediterranean shallow water coral.
Nachshon Siboni
Effect of coal pollution on Mediterranean benthic communities.
Gidon Winters

Photoinhibition in shallow water colonies of Stylophora pistillata as measured in situ. (Co-supervisor Sven Beer )

Noa Levin

Environmental factors influencing the chronic bleaching of the Mediterranean stony coral Oculina patagonica .

Meir Sussman

Fluorescent in situ Hybridization (FISH) reveals the fireworm Hermodice carunculata as a reservoir and a possible vector for the coral pathogen Vibrio shiloi  (Co-supervisor-E. Rosenberg ).

Ofer Ben-Zvi

Deterioration Index (DI): a suggested tool for monitoring reef-coral community health.

(Co-supervisor- A. Abelson).

Omer Polack

Reproductive cycle of Palythoa sp. at Eilat, Red Sea.

(Co-supervisor- Y. Benayahu).

Amy Shlesinger

Nematocysts' toxins of the Mediterranean sea-anemone Aiptasia diaphana and their role in external digestion.

Rachel Armoza
Bio-indicators of stress in scleractinian corals.
Roee Segal

Molecular characteristics of the bleaching phenomenon of the Mediterranean stony coral Oculina patagonica .

Yehonatan Sharon

Ecology of Lessepsian mussels invading the Mediterranean

(Co-supervisor- Y. Benayahu TAU)

Daniel Allen

Community structure of deep (50 m ) scleractinian corals in Eilat, Red Sea .

Ada Alamaru

Trophic biology of Scleractinian planulae

(Co-supervisor- A. Shemesh, Weitzman Institute).

Ido Mizrachi

Sclerochronology of bleached and non-bleached corals.

(Co-supervisor- A. Shemesh, Weitzman Institute).

Maya Weizel

The re-morphogenesis Process in Fungia granulosa: Research and Application

Rafi Yaabetz
Reproductive cycle of a nudibranch.

Omri Bronstein

Morphological and molecular aspects of sea urchins (genus Echinometra)from Okinawa, Zanzibar and Eilat

Ayelet Dadon

Host – symbiont relationship in apo-symbiotic colonies of the Mediterranean coral Oculina patagonica (Co-supervisor M. Fine, Bar Ilan University).

Ran Sulam

Scleractinian coral community in the northern Gulf of Eilat (Red Sea): Effects of a coral disease and a storm


Itzhak Hoskin

Life history strategies of fungiid corals

Gal Eyal Ecology and Taxonomy of mesophotic communities in Israel (Red Sea and Mediterranean)
Elad Mills

Bleaching of the coral Oculina patagonica : role of bacteria in coral health and disease (Co-supervisor- E. Rosenberg)

Ram Barankin
Legislation of Marine Protected Areas in Israel : Mediterranean and Red Sea
Lee Eyal
Reproductive strategies of deep reef ( 60 m depth) corals
Tom Schlesinger
Recruitment of stony corals along the shallow reefs of Eilat
Or Ben-Zvi

Fluorescence in shallow vs. deep water corals

Tal Amit
Microbial populations on corals in shallow and deep (mesophotic) coral populations

Hanna Rapuano

Reproductive strategies of shallow gonochoric corals

Bar Feldman Reproductive strategies of selected mesophotic corals vs. shallow corals

Mila Grinblat

Connectivity between mesophotic corals and shallow corals



Selected Publications



                     Y. Loya and R. Klein "Coral Reefs" (1994).320pp (in Hebrew).

         Ministry of Defence. The publishing House



  Y. Loya and R. Klein (1996). "Coral Reefs" Translated to German

                     "Die Welt der Korallen Encyclopadie der Unterwasserwelt"

                      312 pp Jahr–Verlag GmbH & Co. Hamburg

    Y. Loya and R. Klein (1996) "Coral Reefs" Translated to Dutch-

                    "De Wereld van het koraalrif: Encyclopedie van de

                    onderwatererwereld" 312 pp Jahr–Verl




                    E. Rosenberg and Y. Loya (eds.) (2004).

                 "Coral Health and Disease"

              Springer, Berlin, Heidelberg, New York


Cover page publications

                  (See list of publications for corresponding publication number)


                             64                                                                            110                                                                   138   


                                142                                                                           147                                                                    149                   



                         185                                                            194                                                                          215

                                                     Selected publications (articles)


Y.Loya, L.B. Slobodkin (1971). The coral reefs of Eilat (Gulf of Eilat, Red Sea).

Proc. Zool. Soc. London 28: 117-140.


Y. Loya (1972). Community structure and species diversity of hermatypic corals at Eilat, Red Sea.

Mar. Biol., 13: 100-123.


Y. Loya (1975). Possible effects of water pollution on the community structure  of Red Sea corals.

Mar. Biol., 29: 177-185.


Y. Loya (1976a). The Red Sea coral Stylophora pistillata is an r-strategist. Nature, 259: 478-480.


Y. Loya (1976b). Recolonization of Red Sea corals affected by natural catastrophes and man-made perturbations. Ecology, 57: 278-289.


Y. Loya (1976d). Skeletal regeneration rate in a Red Sea scleractinian coral population.

Nature, 261: 490-491.


Y. Loya (1978). Plotless and transect methods. In: Monographs on Oceanic Methodology.

Coral Reefs: Research Methods. D R. Stoddart and R.E. Johannes (eds.). UNESCO Press, 5: 197-218.


Bradbury R.H., Y. Loya (1978). A heuristic analysis of spatial patterns of  hermatypic corals at Eilat, Red Sea. Amer. Natur., 112: 439-507.


Y. Loya , B. Rinkevich (1979). Abortion effects in corals induced by oil-pollution.

Mar. Ecol. Prog. Ser., 1: 77-80.


Rinkevich B., Y. Loya (1979). The reproduction of the Red Sea coral Stylophora pistillata.

I.Gonads and planulae. Mar. Ecol. Prog. Ser., 2: 133-144.


B. Rinkevich & Y. Loya (1979).The reproduction of the Red Sea coral Stylophora pistillata. II. Synchronization in breeding and seasonality of planulae shedding. Mar. Ecol. Prog. Ser. 2: 145-152.


Loya Y., B. Rinkevich (1980). Effects of oil pollution on coral reef communities.

Mar. Ecol. Prog. Ser., 2: 167-180.   

34. B. Rinkevich, Y. Loya (l983) Intraspecific Competitive Networks in the Red Sea Coral Stylophora pistillata. Coral Reefs 1:161-172

Y.Loya, G. Bull and M. Pichon. (1984). Tumor formations in scleractinian corals.

Helgol. wiss. Meeres., 37: 99-112.

46. B. Rinkevich, Y. Loya (l985). Intraspecific competition in a reef coral: effects on growth and reproduction. Oecologia, 66: 100-105.

L. Muscatine, L.R. McCloskey and Y. Loya (l985).A compare ison of the growth rates of zooxanthellae and animal tissue in the Red Sea coral Stylophora pistillata Fifth Int. Coral Reef Congress, Tahiti, 6:119-123.


Y. Schlesinger, Y. Loya (l985). Coral community reproductive patterns: Red Sea versus the Great Barrier Reef.  Science, 228: 1333-1335.


R. Klein, Y. Loya, G. Gvirtzman, P.S. Isdale and M. Susic (l990).Seasonal rainfall in the Sinai desert during the late Quaternary inferred from fluorescent bands in fossil corals.

Nature, 345: l45-l47.


Y. Loya (l990). Changes in a Red Sea Coral Community Structure: A Long-Term Case History Study.

In: The Earth in Transition: Patterns and Processes of Biotic Impoverishment, G.M. Woodwell (ed.),

Cambridge University Press, pp. 369-384.


B. Lazar, Y. Loya (l99l).Bioerosion of coral reefs - a chemical approach.

Limnol. and Oceanog., 36: 377-383.


N.E. Chadwick, Y. Loya (l992). Migration, habitat use and competition among mobile fungiid corals in the Gulf of Eilat, Red Sea. Mar. Biol. ll4: 6l7-623.


A. Abelson, T. Miloh and Y. Loya (l993). Flow patterns induced by substratum and body morphology of benthic organisms and their role in determining food particle availability.

Limnol. and Oceanog., 38: lll6-ll24.

88. N. Shashar, Y. Cohen, and Y. Loya (1993). Extreme diel fluctuations of oxygen in the diffusive boundary layers surrounding stony corals. Biol. Bull. 185: 455- 461.

A. Abelson, D. Weihs and Y. Loya (1994). Hydrodynamic impedance to settlement of marine propagules

and trailing filament solutions Limnol. and Oceanogr., 39: 164-169.


N. Shashar, T. Feldstein, Y. Cohen and Y. Loya (1994). Nitrogen fixation (acetylene reduction) on a coral reef, Coral Reefs, 13: 171-174.


A. Abelson, Y. Loya (1995). Cross scale patterns of particulate-food acquisition in marine benthic environments. Amer. Natur., 145: 848-854.


A. Lotan, L. Fishman, Y. Loya and E. Zlotkin (1995). Delivery of nematocyst toxin.

Nature, 375: 456-457.

102. O. Mokady, B. Lazar and Y. Loya (1996). Echinoid Bioerosion as a major structuring force of Red Sea coral reefs. Biol. Bull. 190:367-372.

A. Kushmaro, Y. Loya, M.Fine and E. Rosenberg (1996).Bacterial infection and coral bleaching.

Nature, 380: 396.


A. Kushmaro, E. Rosenberg, M. Fine and Y. Loya (1997). Bleaching of the coral Oculina patagoniaby Vibrio AK-1. Mar. Ecol. Prog. Ser.147:159-165.


E. Kramarsky-Winter, M. Fine and Y. Loya (1997). Coral polyp expulsion.

Nature, 387:137


U. Oren, B. Rinkevich and Y. Loya (1997). Oriented intra-colonial transport of 14C labeled materials during coral regeneration. Mar. Ecol. Prog. Ser. 161:117-


Y. Shlesinger, T. L. Goulet & Y. Loya (1998). Reproductive patterns of scleractinian corals in the northern

Red Sea. Mar. Biol. 132:691-701.


U. Oren, I. Brickner and Y. Loya (1998). Prudent sessile feeding by the corallivore snail Coralliophila violacea on coral energy sinks. Proc. Roy. Soc .Biol. Sci., 265:2043-2050.


  L. Stone, A. Huppert, B. Rajagopalan and Y. Loya (1999). Mass coral reef bleaching: a recent outcome of increased El-Nino activity? Ecology Letters 2:325-330.


U. Oren, Y. Benayahu, H. Lubinevsky and Y. Loya (2001). Colony integration during regeneration in the stony coral Favia favus. Ecology, 82: 802-813.


Y. Loya, K. Sakai, K. Yamazato, Y. Nakano, H. Sembali, and R. van Woesik (2001). Coral bleaching: the winners and the losers. Ecology Letters, 4:122-131.


M. Fine, H. Zibrowius and Y. Loya (2001). Oculina patagonica : a non- lessepsian scleractinian coral invading the Mediterranean Sea. Mar. Biol. 138: 1195-1203.


O. Choresh, E. Ron and Y. Loya (2001). The 60-kDa Heat Shock Protein (HSP60) of the sea anemone Anemonia sulcata: a potential early warning system for monitoring environmental changes.

Mar. Biotech. 3: 501-508.

147. M. Fine, Y. Loya (2002). Endolithic algae and coral bleaching.Proc. Roy. Soc. of London - Biological Science, 269: 1205-1210

M. Fine, Y. Loya (2003). Alternate competitive superiority during coral bleaching. Mar. Biol. 142: 989-996.


Y. Loya, E. Kramarsky-Winter (2003). In situ eutrophication caused by fish farms in the northern Gulf of Eilat (Aqaba) is beneficial for its coral reefs: a critique. Mar. Ecol. Prog. Ser. 261:299-303.


Rosenfeld, A. Shemesh and Y. Loya (2003) Implication of water depth on stable isotope composition and skeletal and density banding. Coral Reefs, 22: 337-345.


Y. Loya (2004). The coral reefs of Eilat- past, present and future: Three decades of coral community structure studies. In: Coral Reef Health and Disease; Rosenberg and Loya (Eds).

Springer-Verlag; Berlin Heidelberg, New York. pp. 1-34.


Y. Loya, H. Lubinevsky, M. Rosenfeld and E. Kramarsky-Winter (2004). Nutrient enrichment caused by in situ fish-farms is detrimental to coral reproduction. Mar. Pollut. Bull. 49: 344-353


Y. Loya, M. Rosenfeld and E. Kramarsky-Winter (2005). Nutrient enrichment and coral reproduction: empty vessels make the most sound (response to a critique by B. Rinkevich). Mar. Pollut. Bull. 50: 114-118


Y. Nozawa, Y. Loya (2005). Genetic relationship and maturity state of the allorecognition system affect contact reactions in juvenile scleractinian corals. Mar. Ecol. Prog. Ser. 286:115-123


N. Shenkar, M. Fine and Y. Loya (2005). Size matters: - bleaching dynamics of the coral Oculina patagonica. Mar. Ecol. Prog. Ser. 294: 181-188.


D. Kelman, Y. Kashman, E. Rosenberg, A. Kushmaro and Y. Loya (2006).Antimicrobial activity of Red Sea coralsMar. Biol. 149: 357–363


I. Brickner, U. Frank, U. Oren, and Y. Loya (2006). Energy integration between the solitary polyps of the clonal coral Lobophyllia corymbosa. Jour. of Exp. Biol. 209: 1690-1695


N. Shenkar, M. Fine, E. K. Winter and Y. Loya (2006). Population dynamics of zooxanthella during a bacterial bleaching event. Coral Reefs, 25: 223–227


M. Rosenfeld, A. Shemesh R. Yam and Y. Loya (2006). 18O record of Porites spp. corals during the 1998-bleaching event in Sesoko Island, Okinawa, Japan. Mar. Ecol. Prog. Ser., 314: 127–133


L. Reshef, O. Koren, Y. Loya, I. Zilber-Rosenberg and E. Rosenberg (2006). The Coral probiotic hypothesis, Environ. Microbiol. 8:2068-2073


G. Winters, S. Beer and Y. Loya (2006). In situ measured seasonal variations in Fv /Fm of two common Red Sea corals. Coral Reefs 25:593-598

177. R. Efrony, Y. Loya, E. Bacharach and E. Rosenberg (2007). Phage therapy of coral Disease, Coral Reefs 26: 7-13

T.D. Ainsworth, E. Kramasky–Winter, Y. Loya, O. Hoegh-Guldberg and M. Fine (2007). Coral Disease Diagnostics: What’s between a plague and a band? Appl. Environ. Microb. 73: 981-992


Y. Loya (2007). How to influence environmental decision makers? The case of Eilat (Red Sea) coral reefs. Jour. Exp .Mar. Biol. Ecol. 73: 35-53


O. Choresh, A. Azem and Y. Loya (2007). Over-expression of highly conserved mitochondrial 70-kDa heat-shock protein in the sea anemone Anemonia viridis. Jour. of Thermal Biol. 32: 367–373


A. Zvuloni, Y. Artzy-Randrup, L. Stone, R. van Woesik and Y. Loya (2008). Ecological size-frequency distributions: how to prevent and correct biases in spatial sampling Limno. and Oceanog. : Methods. 6: 144-15


N. Shenkar, Y. Zeldman and Y. Loya (2008). Ascidian recruitment patterns on an artificial reef in Eilat (Red Sea). Biofouling 1-2:119-128.


Harel M, Ben Dov E, Rasoulouniriana D , Siboni N , Kramarsky-Winter E, Loya Y. Barak Z WiesmanZ and Kushmaro A. (2008). A new Thraustochytrid, strain Fng1, isolated from the surface-mucus of the hermatypic coral Fungia granulosa FEMS Microbiol. Ecol. 1-10.


Assaf Zvuloni, Rachel Armoza-Zvuloni1 and Yossi Loya (2008). Structural deformation and growth inhibition of branching corals associated with the vermatid gastropod Dendropoma maxima

Mar. Ecol. Prog. Ser 363:103-108.


Y. Loya and K. Sakai (2008). Bidirectional sex change in mushroom corals. 

Proc. Roy. Soc. Biol. B 275:2335-3343.


N. Shenkar Y. Loya (2008). Ecology and systematics of the ascidian fauna in the Gulf of Eilat (Aqaba).

In “Aqaba-Eilat, the Improbable Gulf. Environment, Biodiversity and Preservation” Ed. F.D Por, Magnes, Jerusalem. pp 197-208.


N. Shenkar, O. Bronstein and Y. Loya (2008) population dynamics of a coral reef ascidian in a deteriorating environment. Mar. Ecol. Prog. Ser., 367:163-171.


Eugene Rosenberg , Ariel Kushmaro, Esti Kramarsky-Winter, Hudi Banin, and Yossi Loya (2008). The role of microorganisms in coral bleaching. Inter. Soc. Micr. Ecol. (ISME ) 3: 139-146.


N. Shenkar Y. Loya (200 8 ). The solitary ascidian Herdmania momus : native (Red Sea) versus

non-indigenous (Mediterranean) populations . Biol. Inv., 10: 1431-1439.


A. Zvuloni, Y. Artzy-Randrup, L. Stone, E. Kramarsky-Winter, R. Barkan and Y. Loya (2009).

Spatio-temporal transmission patterns of black-band disease in a coral community.

PLoS ONE 4:1-10.

Ami Schlesinger, Eliahu Zlotkin, Esti Kramarsky-Winter and Y. Loya (2009). Cnidarian internal stinging mechanism. Proc. Roy. Soc. Biol. B 276:1063-1067.


Gidon Winters, Sven Beer and Yossi Loya (2009). Spatial and temporal photoacclimation of Stylophora pistillata : zooxanthella size, pigmentation, location and clade. Mar. Ecol. Prog. Ser. 384:107-119.

193. A. Alamaru, R. Yam, A. Shemesh and Y. Loya (2009). Trophic Biology of Coral Larvae: Evidence from Stable Isotope Analysis Mar. Ecol. Prog. Ser. 383:85-94.

A. Schlesinger, E. Kramarsky-Winter and Y. Loya (2009). Active nematocyst isolation via nudibranchs

Mar. Biotechnol. 11:441–444.


A. Alamaru, O.Bronstein, G.Dishon and Y.Loya (2009). Opportunistic feeding by the fungiid coral Fungia scruposa on the moon jellyfish Aurelia aurita Coral reefs 28:865


E. Kramarsky-Winter, C. A. Downs, A, Downs and Y. Loya (2009). Cellular responses in the coral Stylophora pistillata exposed to eutrophication from fish mariculture . Evol. Ecol. Res. 11: 1–21.


M.Vizel, E.Kramarsky-Winter and Y.Loya (2009). Mushroom coral regeneration from a detached stalk.

Coral reefs 28:939


G.Tsagkogeorga, X. Turon, R. Hopcroft, M.-Ka Tilak, T. Feldstein, N. Shenkar Y. Loya , D. Huchon, E. Douzery and F. Delsuc (2009) An updated 18S rRNA phylogeny of tunicates based on mixture and secondary structure models BMC Evolutionary Biology.


N. Shenkar Y. Loya (2009). Non-indigenous ascidians (Phylum: Chordata, Subphylum: Tunicata) along the Mediterranean coast of  Israel Mar. Biod. Rec. doi:10.1017/S1755267209990753;Vol.2;e166;2009


A. Schlesinger, R. Goldshmid, M. G. Hadfield, E. Kramarsky-Winter and Y. Loya (2009). Laboratory culture of the aeolid nudibranch Spurilla neapolitana (Mollusca, Opisthobranchia): life history aspects.

Mar Biol 156: 753-761.


C. A. Downs, E. Kramarsky-Winter, J. Martinez, A. Kushmaro, C.M. Woodley, Y. Loya , G. K. Ostrander (2009). Symbiophagy as a Cellular Mechanism for Coral Bleaching. Autophagy 5: 211-216.


D. Kelman, Y. Kashman, R. Hill, E. Rosenberg and Y. Loya (2009). Chemical warfare in the sea: The search for antibiotics from Red Sea corals and sponges Pure Appl. Chem. 6:1113-1121.


C.A. Downs, E. Kramarsky-Winter, C. M. Woodley, A. Downs, G. Winters, Y. Loya , Gary K. Ostrander (2009). Cellular pathology and histopathology of hypo-salinity exposure on the coral Stylophora pistillata. Science Tot. Envir   407: 4838-4851.


A. Alamaru., Y. Loya, E. Brokovich , R. Yam and A. Shemesh (2009) Carbon and nitrogen utilization in two species of Red Sea corals along a depth gradient; insights from stable isotope analysis of lipids.

Geoch. and Cosmoch. Acta 73:5333-5342.


G. Winters, R. Holzman, A. Blekhman, S. Beer and Y. Loya (2009). Photographic assessment of coral chlorophyll contents: Implications for ecophysiological studies and coral monitoring.

Jour. Exp. Mar. Biol. Ecol . 380: 25-35.

207. L. Arotsker, N. Siboni, E. Ben-Dov, E. Kramarsky-Winter, Y. Loya and A.
Kushmaro (2009). Vibrio sp. as a potentially important member of the Black Band Disease (BBD) consortium in Favia sp. corals. FEMS Microbiol Ecol 70: 515-524
208. T. R. Singh, G. Tsagkogeorga, F. Delsuc, S. Blanquart, N. Shenkar , Y. Loya , E.J.P Douzery and D. Huchon Tunicate mitogenomics and phylogenetics: peculiarities of the  Herdmania momus  mitochondrial genome and support for the new chordate phylogeny. BMC Genomics  10 : 534doi:10.1186/1471-2164-10-534s.

Y. Loya , K. Sakai and A. Heyward (2009). Reproductive patterns of fungiid corals in Okinawa , Japan . Galaxea Jour. of Coral Reef Stud. 11: 119-129.


A. Zvuloni, R. van Woesik and Y. Loya (2010) Diversity Partitioning of Stony Corals

Across Multiple Spatial Scales Around Zanzibar Island, Tanzania.

PLoS ONE5(3):e9941.doi:10.1371/journal.pone.0009941.


N. Siboni, D. Rasoulouniriana, E. Ben-Dov, E. Kramarsky-Winter, A. Sivan, Y. Loya , O. H. Guldberg and A. Kushmaro (2010) . Stramenopile Microorganisms Associated with the Massive Coral Favia sp J. Eukaryot. Microbiol . pp.1–9


A. Schlesinger, E. Kramarsky-Winter, H. Rosenfeld, R. Armoza-Zvoloni and Y. Loya (2010). Sexual plasticity and self-fertilization in the sea anemone Aiptasia diaphana. PLoSONE 5(7):e11874. doi:10.1371/journal.pone.0011874


I. Brickner , Y. Loya and Y. Achituv (2010). Diverse life strategies in two coral- inhabiting barnacles occupying the same host (Cyphastrea chalcidicum), in the northern Gulf of Eilat . Jour. Exp .Mar. Biol. Ecol. 392:220-227


I. Mizrachi ,Y. Loya, M. Rosenfeld, E. Kramarski- Winter, R. Yam and A. Shemesh (2010). The stable isotope composition of newly formed skeleton in the stony coral Porites spp.

Geochimica et Cosmochimica Acta 74:7021–7030

215. M.Vizel, Y. Loya, C. A. Downs and E. Kramarsky-Winter (2010). A novel method for coral explant culture and micropropagation. Mar. Biotechnol. DOI10.1007/s10126-010-9313-z
216. Wild et al (2011). Climate change impedes scleractinian corals as primary reef ecosystem engineers. Marine and Freshwater Research 62:205–215.
217. O. Bronstein and Y. Loya (2011) Day time spawning of Porites rus on the coral reefs of Chumbe Island in Zanzibar , Western Indian Ocean (WIO). Coral Reefs   30:44

R. Armoza-Zvuloni , E. Kramarsky-Winter & Y. Loya (2011). Repeated bleaching events may result in high tolerance and notable gametogenesis in stony corals: Oculina patagonica as a model.

Mar. Ecol. Prog. Ser . 426:149-159


C. Mora, O. Aburto-Oropeza, A. Ayala-Bocos S. Banks … Y. Loya, et al. (2011). Global human foot print on the linkage between biodiversity and ecosystem functioning in reef fishes.

PloSBio 19(4):e1000606.doi:10.1371/journal.pbio.1000606220

220.   G. Eyal, L. Eyal-Shaham and Y. Loya (2011). "Teeth-anchorage": sleeping behavior of a Red-Sea filefish on a branching coral Coral Reefs 30: 707

R. van Woesik, K. Sakai, A. Ganase & Y. Loya (2011). Revisiting the winners and the losers a decade a decade after coral bleaching Mar. Ecol. Prog. Ser . 434:67-76


O. Polak, Y. Loya , I. Brickner and Y. Benayahu (2011). The widely distributed indo-pacific zooanthid Palythoa tuberculosa: a sexually conservative strategist. Bull. Mar. Sci 87:605-621

223. A. Atad, A. Zvuloni, Y. Loya and E. Rosenberg (2012). Phage therapy of the white plague- Like disease of Favia favus in the Red Sea. Coarl Reefs 31: 665-670.
224. Armoza-Zvuloni, R., E. Kramarsky-Winter, H. Rosenfeld, L.S Shore, D Sharon & Y. Loya (2012). Reproductive characteristics and steroid levels in the scleractinian coral Oculina patagonica inhabiting contaminated sites along the Israelí Mediterranean coast. Mar. Pollut. Bull. 64:1556-1563
225. MC. Ojimi, Y. Loya and M. Hidaka (2012). Sperm of the solitary coral Ctenactis echinata exhibit a longer telomere than that of somatic tissue. Zoological Studies 51: 1475-1480.
226. Paramasivam N., E. Ben-Dov, L. Arotsker, E.Winter, A. Zvuloni, Y. Loya and A. Kushmar (2013). Bacterial consortium of Millepora dichotoma exhibiting unusual multifocal lesion event in the Gulf of Eilat, Red Sea. Microb. Ecol. 65:50-59.
227. Shashank Keshavmurthy et al. (2013). DNA barcoding reveals the coral “laboratory-rat”, Stylophora pistillata encompasses multiple identities. Scientific Reports 3: DOI: 10.1038/srep01520 Published: March 22, 2013.
228. Bronstein O. & Y. Loya (2013). The taxonomy and phylogeny of echinometra (camarodonta: echinometridae) from the red sea and Western Indian ocean. PLoS ONE Volume: 8 Issue: 10 Pages: e77374 DOI: 10.1371/journal.pone.0077374
229. Mills E., K. Shechtman, Y. Loya & E. Rosenberg (2013). Bacteria cause and prevent bleaching of the coral Oculina patagonica. Mar. Ecol. Prog. Ser 48: 155-162
230. Kramarsky-Winter E, L. Arotsker, D. Rasoulouniriana, N. Siboni, Y. Loya and A. Kushmaro (2013). The Possible Role of Cyanobacterial Filaments in Coral Black- Band Disease PathologyInvert. Microb. DOI 10.1007/s00248-013-0309 published on line 20.10. 2013
231. Downs CA, McDougall KE, Woodley CM, Fauth JE, Richmond RH, Kushmaro A, Gibb AS, Loya Y, Ostrander GK and E. Kramarsky-Winter (2013). Heat-Stress and Light-Stress Induce Different Cellular Pathologies in the Symbiotic Dinoflagellate during Coral Bleaching. PLoS ONE 8(12): e77173. doi:10.1371/journal.pone.0077173

Downs C.A., Kramarsky-Winter E., Fauth J. E., Segal R., Bronstein O., Jeger, R., Lichtenfeld, Y., Woodley M., Pennington P., Kushmaro A. & Y. Loya (2013). Toxicological effects of the sunscreen UV filter, benzophenone-2, on planulae and in vitro cells of the coral Stylophora pistillata Ecotoxicology.

233. Armoza- Zvuloni R., E. Kramarsky-Winter, Y. Loya, A. Schlesinger & H. Rosenfeld (2013). A unique sexual system in a sea anemone provides first evidence of trioecy in the animal kingdom. Biology of Reproduction, DOI:10.1095/biolreprod.113.114116

Bronstein O. & Y. Loya (2014). Echinoid community structure and rates of herbivory and bioerosion on exposed and sheltered reefs. Jour. Exp .Mar. Biol. Ecol .456: 8-17.

235. Loya Y., A. Genin A., Al-Zibdah M., Naumann MS. & C.Wild (2014). Reviewing the status
of coral reef ecology of the Red Sea - the need to consider key topics and relevant research.
Coral Reefs, 33:1179–1180
236. Bronstein O. & Y. Loya (2014). Photoperiod, temperature and food availability as drivers
of the annual reproductive cycle of the sea urchin Echinometra sp. from on the Gulf of Aqaba
(Red Sea) Coral Reefs, DOI 10.1007/s00338-014-1209-3.
237. Bronstein O. & Y. Loya (2014). Photoperiod, temperature and food availability as drivers
of the annual reproductive cycle of the sea urchin Echinometra sp. from on the Gulf of Aqaba
(Red Sea) Coral Reefs, DOI 10.1007/s00338-014-1209-3.
238. Adi Lavy ,Gal Eyal, Benjamin Neal, Ray, Yossi Loya and Micha Ilan (2015).
A quick, easy, and non-intrusive method for underwater volume and surface area evaluation of
benthic organisms by 3D computer modeling (Methods in Ecology and Evolution; accepted
for publication)


Full CV                 

 Academic and Professional experience

      Department/ Division


Israeli Academy of Sciences and Humanities
Science Division



Center of Excellence, James Cook University, Townsville, Australia
Coral Reef Ecology
Visiting Professor
James Cook University, Dept. of Marine Biology, Townsville, Australia
Coral Reef Ecology



Tel Aviv University

The Raynor Chair for Environmental
Conservation Research



95, 97, 99, 04,

05, 07, 08, 09, 10, 12, 14 (summers)

University of the Ryukyus,
Okinawa, Japan

Sesoko Tropical Biosphere

Research Center

Visiting Professor


Tel Aviv University
The Porter Super-Center for Ecological and Environmental Studies



Tel Aviv University
Faculty of Life Sciences
Tel Aviv University
Tel Aviv University
Full Professor
Tel Aviv University
Associate Professor



Scripps Inst. of Oceanography

La Jolla, San Diego USA



Visiting Professor

“Coral Reefs” Journal Springer-Verlag
(Biology section)



Australian Institute of Marine Sciences (AIMS) and James Cook University, Australia


Marine Ecology

Visiting Professor
(Senior Queen Elizabeth Fellow)
Tel Aviv University
Senior lecturer
Tel Aviv University



Woods Hole Oceanographic Institute, Woods Hole, Mass.





State University of New York at Stony Brook,

L.I., N.Y.


Senior teaching

Univ. of Michigan,

Ann Arbor

Teaching assistant
Tel Aviv University
Teaching assistant


International Committees

Advisory and Administrative duties

2012- present

Advisory Board: Batsheva de Rothschild Fund for the Advancement     of Science in Israel.

2011- present
Chairman- Israeli Scientific Committee Oceanic Research (SCOR).
2011- present
Chairman- MA'RAG Israel Academy of Sciences and Humanities.
2006- present
Scientific Board of the Australian Research Council (ARC) Centre of Excellence on Coral Reef Research.
Central Nomination Committee Tel Aviv University.
Advisory and Review Board of the Research Institute of the Subtropics, Okinawa, Japan (Coral bleaching project).
   Co - Chairman  

International Targeted Working Group on indicators of coral bleaching under the auspices of GEF and the World Bank .

The Scientific Review Board- Coral bleaching Project,
Research Institute for the Subtropics (RSI), Okinawa, Japan.
The Board of Directors IUI, Eilat.
Academic Planning Committee, Tel Aviv University.
The Board of Directors, Inter-University Institute
of Marine Science (IUI), Eilat.
Israel Science Foundation (Ecology committee).
Israeli Ministry of Science (Germany-Israel Marine Biology Program).
Israeli Man and the Biosphere Committee. (MAB)- UNESCO.
Board of Trustees, Tel Aviv University.
Board of Governors, Tel Aviv University.
University Central Committee, Tel Aviv University.
Faculty of Life Sciences, Tel Aviv University.
The Israeli Scientific Delegation to the UN Conference on
Environmental Development (UNCED) Rio-de Janeiro, Brazil.

Department of Zoology, Tel Aviv University.

The Scientific Review Board, Oil Spill Project-
Smithsonian Tropical Research Institute (STRI, Panama).
“Coral Reefs”, Springer-Verlag, (Biology Section).
Council Member
International Society for Reef Studies (ISRS).





International Association of Biological Oceanography (IABO)

organizing the following coral reef symposia:
-3d International Symposium on Coral Reefs: Miami, Florida, 1977
-4th International Symposium on Coral Reefs: Philippines, May 1981.
-5th International Symposium on Coral Reefs: Tahiti, June l985.
-6th International Symposium on Coral Reefs: Australia, August l988.
-7th International Symposium on Coral Reefs: Guam, June l992.

Scientific advisory board for the Nature Protection
Authority, Israel
Israel National Academy of Science Foundation.



The editorial board of the journals:
Marine Ecology Progress Series(MEPS,1979-1984);
Marine Biology (1986-1990); Marine pollution Bulletin (2002-present)
The Zoological Society of Israel.


Scientific Committee on Oceanic Research (SCOR, UNESCO),
Working group on: “Quantitative studies on coral reefs”






   The Darwin Medal

 (Award conferred once every 4 years)

Awarded by the International Society for Reef Studies (ISRS) for life contribution to coral reef research.

Award conferred during the 9th International coral Reef Symposium Bali, Indonesia.


The Landau Prize

(together with Eugene Rosenberg, Tel Aviv University)

Awarded by Mifaal Hapais in the category of Life Sciences for original outstanding research contributionto the field of Ecology and Environmental Quality.


2013-2016 Middle East Regional Cooperation (Program: U.S. Agency for International
Development, Bureau for the Middle East (MERC-AID)
: Developing novel
methodologies for preservation and maintenance of coral biodiversity.
2012-2016 Israel Science Foundation (ISF) Sex-allocation and sex change in mushroom scleractinian corals: a long term study.
2012-2013 Israel Taxonomy initiative (ITI): Biodiversity of Mesophotic Scleractinian corals in
the Gulf of Eilat/Aqaba.

2012-2013 Israel Taxonomy initiative (ITI): Taxonomy & molecular systematics of the
Ctenophore fauna along the coasts of Israel (PI in collaboration with Ada Alamaru,
and Dorothee Huchon, TAU)


Israel Science Foundation (ISF)- Etiology of Black Band Disease (BBD) in the Red Sea. (PI in collaboration with E. Rosenberg, TAU)


Marie Curie European Research Training Network (with M. Ilan, TAU)

2004- 2008

Israel Science Foundation (ISF): "The isotopic composition of Eilat's corals:

basic aspects of signals buildup and tracing anthropogenic stress.

(Co-PI in collaboration with A. Shemesh, the Weizman Institute).

2004- 2007

The Porter School of Environmental Studies in collaboration with the Italian Ministry

of the Environment: "Artificial Marine Structures (AMS): Multifunctional Tool for Research and Environmental Management in the Mediterranean and Red Sea (MED- RED). (PI in collaboration with Y. Benayahu and A. Abelson).

2003- 2007

Israel Science Foundation (ISF): “An integrative approach of studying bacterial coral bleaching in the coral reefs of Eilat". (in collaboration with E. Rosenberg TAU).


2005- 2010

GEF/The World Bank/UNESCO/IOC : International Targeted Working Group of experts on "indicators of coral bleaching". Co-Chairman with Professor Ove Hoegh Guldberg (Chairman) of the group, including 15 scientists from USA, England, Australia, Kenya, Israel, and Mexico.The group meets annually and works together 2-3 weeks

at four reef sites: Heron Island (Great Barrier Reef, Australia), Puerto Morelos (Mexico), Philippines and Zanzibar.


Israel Science Foundation (ISF):
“Alternative feeding mechanism in corals: bacterial aggregate “gardening”.
(in collaboration with A. Kushmaro, Ben Gurion Univ.)

MERC-Bi-national Research and Monitoring Program (Israel -Jordan) for

the Red Sea Marine Peace Park, Aqaba, Jordan


Israeli Ministry of Science: “Establishment of a National Infrastructure
Laboratory: Center for High Throughput Screening (HTS) for Novel Bioactive Compounds at Tel Aviv University”-

(Director and Coordinator of a research project involving 22 scientists

from 6 academic and research institutions in Israel).


Israeli Ministry of Science:
Novel Bioactive Compounds from Marine Invertebrates and their symbionts
(in collaboration with Dr. M. Ilan, TAU).


The German Ministry of Education, Science, Research and Technology
(B.M.B.F.):- Red Sea Program-"Scleractinian corals as environmental

recorders of the Red Sea”.


U.S.-Israel Bi-National Science Foundation (BSF):
“Coral bleaching by bacteria”.(In collaboration with E. Rosenberg, TAU).
Israel Science Foundation (ISF):
”Bioerosion of coral reefs a multidisciplinary approach”.
(in collaboration with B. Lazar, the Hebrew University)


The German-Israeli Foundation for Scientific Research and

Development (G.I.F): “Patterns of stable isotope fractionation during mineralizationprocesses in corals: Environmental implications”.

Israel Academy of Sciences-Basic Research Foundation:
“Bioerosion of coral reefs by sponges, sea-urchins and bivalves”.


US National Institute of Health (NIH):
“Novel natural products from the sea as potential anti-AIDS drugs”.-
(in collaboration with Prof. Y. Kashman, School of Chemistry
and Prof. A. Hizi, Medical School, TAU).
Office of Environmental Quality:
“Medusa blooms in the Mediterranean Sea: Ecological aspects”.
National Coal Company: “Effects of coal pollution on coral reefs”.

FAO Mediterranean Action Plan (MED POL) - “Swarming of jellyfish

along the Mediterranean Coast of Israel: An environmental approach”.


National Council for Research and Development -

Joint German-Israeli Program: “A new method to assess

indicators of environmental stress and possible stabilization or

degradation of toxicants in coral reefs”.


Harbor Branch Oceanographic Institution, Sea Pharm Project:
“Bioactive materials derived from soft corals and sponges from the

Red Sea”. (in collaboration with Prof. Y. Kashman, TAU).

U.S.-Israel Bi-national Science Foundation (BSF):
“Larval ecology of boring bivalves in living corals”.
Israeli Academy of Sciences- Basic Research Foundation:
“Reproductive strategies of Red Sea scleractinian corals”.
The Society for Nature Protection: “Massive predation of
scleractinian corals at Eilat caused by a predatory gastropod”.
The Fund for Basic Research, Tel Aviv University:
“Ecology of boring bivalves in living stony corals”.
U.S.-Israel Bi-national Science Foundation (BSF):
“Isolation and identification of New Marine Natural Products”.
(in collaboration with Prof. Y. Kashman, TAU).


Israeli Academy of Sciences- Basic Research Foundation:
“The variation in the chemical composition of soft-corals and

sponges as a function of the place of collection”.

(in collaboration with Prof. Y. Kashman,TAU).

Senior Queen Elizabeth Fellowship:
Australian Institute of Marine Science, Cape Ferguson, Australia.
Ministry for Commerce and Industry:
“Biologically active products from marine organisms”.
(in collaboration with Prof. Y. Kashman, Tel- Aviv University).
National Council for Research and Development:
“Experiments on the effect of oil pollution on hermatypic corals”.
U.S.-Israel Bi-National Science Foundation:(BSF):
“Growth and development of reef corals.”
National Council for Research and Development:
“The effect of pollutants on coral reef communities”.
NOAA - Manned Undersea Science and Technology Program:
“External distribution of energy fixed by reef corals at Puerto-Rico”.
Two weeks underwater expedition using underwater laboratory-PRINUL
(in collaboration with Professors. S. Richman and Dr. L. McCloskey).


          Membership in professional societies            

The Israeli Association for Aquatic Sciences

Ecological Society of Israel

Zoological Society of Israel
  Ecological Society of America 

International Society for Reef Studies



Year                                                 Meeting
2014 Second International Mesophotic Coral Reef Workshop (MCEIsrael), Oct. 26-31, Inter-University Institute (IUI), Eilat, Israel (Chairman and organizer of workshop).
2013 Australian Research Council (ARC) Center of Excellence Symposium, Townsville, Australia, 9-12 October 2013 Coral Reefs in the 21st Century
Plenary Lecture: Reproductive Strategies of Fungiid Corals

Eighth International Conference on Coelenterate Biology (ICCB8), Dec 1-7 Eilat, Israel

Plenary Lecture: Reproductive Strategies of Fungiid Corals in the Coral Reefs of Okinawa, Eilat and Aqaba

2012 Amakusa Biodiversity Symposium: Biodiversity in Changing Coastal Waters of Tropical and Subtropical Asia November 29-Dec.4; Amakusa Japan
Lecture: Fungia fungites: a unique sex changing brooder among the Fungiidae.

The 12th International Coral Reef Symposium 9-13 July Cairns, Queensland, Australia.

Lecture: The solitary coral Fungia fungites is a protogynous sequential hermaphrodite Brooder.

2011 Annual Zoological Society of Israel 25th Dec. Tel Aviv University
Recipient of honorary membership of the Zoological Society of Israel
Plenary lecture: Reproductive strategies in mushroom stony corals
2011 ARC Center of Excellence symposium: Coral Reefs: Coast to Coast symposium. 14-16 October. Fremantle, Perth Western Australian Maritime Museum

ARC Center of Excellence symposium: "Coral reefs in a changing environment", 7-8 October, Canberra , Australia

Key note address : "Multiple sex change in mushroom corals"


11th International Symposium on Spermatology; Satellite Symposium, 30 th June, Motobu, Okinawa , Japan .

Lecture: "Fungiid corals: ideal model organisms to study evolution of coral reproductive traits"


The 12th Japanese Coral Reef Society meeting. Memorial lecture:the late Professor Kiyoshi Yamazato. 28-29 November, Okinawa , Japan .

Plenary Lecture: "Reproductive Patterns of Fungiid Corals in Okinawa , Japan ".
2009 Organizer of a joint workshop of the Australian Research Council (ARC) Center of Excellence(CoE) for Coral Reef Studies and the Interuniversity Institute for Marine Sciences in Eilat (IUI): "Coral reefs of the Indo-Pacific in an era of global change" held at IUI, Eilat, Israel, 5-6 October.
2009 The 12th Japanese Coral Reef Society meeting. Plenary memorial lecture the late Professor Kiyoshi Yamazato, 28-29 November, Okinawa, Japan.
Lecture: "Reproductive Patterns of Fungiid Corals in Okinawa, Japan".
2009 GEF/World Bank working group on coral bleaching Workshop
May 15-30, Heron Island, Australia
2009 International Okinawa Churaumi Aquarium Symposium on
"Reproduction of marine organisms" Feb. 21-23, Okinawa, Japan
Lecture: Repeated sex change in mushroom corals
2008 First International Congress Documenting, Analyzing and Managing
Biodiversity in the Middle East. October 20-23, Aqaba, Jordan
Lecture: The Coral Reefs of Eilat-past present and future.
2008  The 11th International Coral Reef Symposium, June 7-11, Miami,
Fort Lauderdale, USA
Lecture: Bidirectional sex change in fungiid corals.
2008  IOC-GEF/World Bank/UNESCO working group on coral bleaching Workshop
February 12-16, Amsterdam, the Netherlands.
2007 GEF working group on coral bleaching Workshop, Nov. 20-25, Miami, Florida
2007 ARC Centre of Excellence- second scientific annual board meeting,
February, Sydney, Australia.
Lecture: The Coral Reefs of Eilat: Three decades of coral community structure studies.


ISRS (International Society for Reef Studies) European Meeting Bremen,

Germany Sept 19-22
Lecture: Net pen fish farming and coral reefs: An unhappy marriage.

2006 Palau Coral reef workshop, Koror 25.6. - 8.7.
Lecture: Fish net pen mariculture and the coral reefs of Eilat: a sad story.

ARC Centre of Excellence- first scientific annual board meeting,

February 17-20, Townsville, Australia.


IOC-GEF/World Bank working group on coral bleaching Workshop,

7-11 April,  Paris .


World Bank/ UNESCO/IOC-GEF International targeted Group of Experts fourth workshop on indicators of coral bleaching, 10-29 May, Puerto Morelos , Mexico.


Peer Review of the Department of Ecology and Evolutionary Biology,

13-16 April, Ravenna , Italy .


Ilanit Congress of the Federation of the Israel Societies for Experimental Biology (FISEB ) February 7-10 Eilat.

Invited Plenary Lecture: The Coral Reefs of Eilat: Three decades of coral community structure studies.


The World Bank/ /IOC-GEF International targeted Group of Experts third   

workshop on indicators of coral bleaching; 8-16 January, Puerto Morelos, Mexico .


IOC-GEF/World Bank working group on coral bleaching Workshop,

19-22  October, Washington DC


Census of Marine Life (Coml) Coral Reef Initiative Working Group:

Quantitative   methodologies for assessing coral community bleaching ,

14-16 October Washington DC


The 10th International Coral Reef Symposium , Okinawa , Japan .

Lecture: How to kill a coral reef?


International Coral Ecotoxicology and Health Workshop: Bermuda Biological Station for Research Invited lecture: The coral reefs of Eilat - past, present and future: Three decades of coral community structure studies.


The Red Sea Marine Peace Park International Symposium Integration of Marine Science and resource Management Aqaba 2-4 December 2003

Workshop on “Coral Health and Diseases”, Eilat, Israel.
Invited lecture: The coral reefs of Eilat: 35 years of monitoring their coral
community structure
Marine Molecular Biotechnology and Biodiversity Meeting; Wendelsheim, Germany.
Plenary lecture: Biodiversity of the Red Sea coral reefs: A unique source for new natural products.

 The World Bank/UNESCO/IOC International Working Group on indicators of coral bleaching:- Workshop on Coral Bleaching: Biological Early Warning Systems, Paris, France.

International Review Board Meeting of the Coral Bleaching Project.
Research Institute of the Subtropics (RIS), Tokyo, Japan.
The World Bank/UNESCO/IOC International Targeted Group of Experts
on "indicators of coral bleaching".
Second Field-Workshop of the Working Group, Puerto Morelos, Mexico.

Workshop on “Biological considerations associated with current geochemical approaches using stony corals as proxies for the reconstruction of different aspects of past climates” Woods Hole Oceanographic Institution, Cape Cod, Mass.
Invited lecture: “Using stony corals as proxies for the reconstruction of Red

Sea past climates”.

Sixth EU Framework Programme of the European Community for research
technological development and demonstration activities; Network of Excellence- Marine Molecular Biotechnology.
Invited lecture: Bioactive Compounds from Red Sea Marine Organisms.
The World Bank/UNESCO/IOC International Targeted Group of Experts on
"indicators of coral bleaching". First Field-Workshop of the Working Group;
Heron Island, The Great Barrier Reef, Australia.
The International Society of Reef Studies European Coral Reef Meeting;
Cambridge, England
Lecture: Bleaching of Mediterranean corals is caused by a bacterial pathogen

The World Bank/UNESCO/IOC Workshop on Indicators of Coral Stress, Paris, France.

Initiation of the International Working Group of Experts.
Lecture: Coral bleaching: the winners and the losers..

The 9th International Coral Reef Symposium, Bali, Indonesia. Awarded the
Darwin Medal (Year 2000) for life time contribution to coral reef research.
Plenary lecture: Homage to Stylophora pistillata: a significant coral species
in reef research.

Japan Marine Science and Technology (JAMSTEC) International Coral reef
Symposium- Coral reef biodiversity and health as indicators of environmental change, Tokyo, Japan.                                                           Invited lecture: Regeneration processes in scleractinian corals.

Expert workshop on coral bleaching: Convention on Biological Diversity
Manila, Philippines.
Invited lecture: Devastating Coral bleaching in Okinawa, Japan.
The Third EuroMab Biosphere Reserves Coordinators’ Meeting Ilomantsi, Finland.
Lecture: The Red Sea Peace Park as a Trans-boundary Marine Biosphere Reserve.

Hawaii Coral Reef Monitoring Workshop-A Tool for Management. Univ. of Hawaii
Plenary lecture: The Red Sea Peace Park Coral Reefs Benthic Communities: Ecology

and Biology Monitoring Program.

Meeting of the Red Sea Program of Marine Sciences, Bremen, Germany.
Lecture: Scleractinian corals as environmental recorders of global change.
Annual meeting of the Society for Integrative and Comparative Biology; Boston, USA.
Lecture: Bleaching of corals caused by bacteria.
Workshop of the Bi-national Research and Monitoring Program (Israel –Jordan)- (MERC) for the Red Sea Marine Peace Park:–Discussion Leader.
Aqaba, Jordan.
Biotic recoveries from mass extinctions; Panel member.
Academy of Sciences, Prague, Czech Republic
Eighth International Coral Reef Symposium, Panama City, Panama.
Lecture: Predation of corals by the predatory snail Drupella cornus.

18th Pacific Science Congress: Population, Resources and Environment Prospects;

Beijing, China.
Lecture: Stony corals as environmental recorders.

International Coral Reef Initiative Workshop; Dumaguete City, Philippines.
Israeli representative discussing quantitative methods for monitoring coral reefs.
Conference of the Red Sea Program (RSP) on Marine Sciences, Dahab, Egypt.
Lecture: The use of scleractinian corals as environmental recorders of the Red Sea.
UNESCO 28th General Meeting:-Scientific Representative of the Israeli delegation proposing to declare the Gulf of Aqaba (Eilat) as a Biosphere Reserve.

Conference on Global Aspects of Coral Reefs: Health, Hazards and History,

Univ. of Miami, USA.
Plenary lecture: Long-term changes in coral community structure at Eilat, Red Sea

Conference on Middle East Multilateral Talks, UCLA, Los Angeles, USA.
Invited lecture: Development and protection of the Gulf of Aqaba.
International Symposium on Biodiversity and Adaptive Strategies of Coral Reef Organisms, Okinawa, Japan. Plenary lecture: Massive predation of scleractinian corals at Eilat caused by the predatory gastropod Drupella cornus.

Status seminar of the German-Israeli Co-operation in Environmental Research; Ploen, Germany.
Lecture: A new method to assess indicators of environmental stress and possible
stabilization of toxicants in the environment.

Seventh International Coral Reef Symposium, Guam,
Lectures: l. Settlement, metamorphosis and bioerosion rate of the boring bivalve Lithophaga lessepsiana.
2. Depth-dependent timing of density band formation in scleractinian corals
at the coral reefs of Eilat, Red Sea, Israel.

Workshop on oil pollution effects on the coral reefs of Panama; Meeting of Scientific Review Board, Phase 3.
Smithsonian Tropical Research Institute, Panama


Workshop on coral bleaching, Coral Reef Ecosystems and Global Climate Change. Miami, Florida.
Invited lecture: Climate change in Sinai Desert during the late Quaternary inferred
from fluorescent bands in fossil corals.


Fifth International Congress of Ecology, Yokohama, Japan.
Invited lecture: Coral host specificity between the date mussel Lithophaga lessepsiana

and the coral Stylophora pistillata.


Workshop on the oil pollution effects on the coral reefs of Panama; Meeting of the Scientific Review Board, Phase 2.
Smithsonian Tropical Research Institute, Panama

Annual meeting of the International Society for Reef Studies, Marseilles, France.
Invited lecture: Bioerosion of coral reefs.
Fifth International Congress of Invertebrate Reproduction, Nagoya, Japan.
Lecture: Resource allocation between growth and reproduction in corals and
boring bivalves.
Fifth International Conference on Coelenterate Biology, Southampton, England.
Lecture: Larval and postlarval recruitment in the broadcasting scleractinians
Favia favus and Platygyra lamellina.
Workshop on the long term assessment of the oil-spill at Bahia Las Minas, Panama.
Washington, DC., USA.
Sixth International Coral Reef Symposium, Townsville, Australia.
Lecture: First sclerochronological record of hermatypic corals from the Red Sea.

Regional Research Workshop and International Symposium on the Conservation and Management of Coral Reef and Mangrove Ecosystems, Okinawa, Japan.
Invited lecture: Effects of man-made versus natural disturbances on coral reefs.


Workshop on oil pollution effects on the coral reefs of Panama; Meeting of

Scientific Review Board, Phase 1.
Smithsonian Tropical Research Institute, Panama.

Twenty-second European Marine Biology Symposium, Barcelona, Spain.
Plenary lecture: Pollution effects of coral reef communities.

The Woods Hole Research Center: Conference on Biotic Impoverishment,
Woods Hole, USA.
Plenary lecture: Changes in a Red Sea coral community under chronic oil pollution:
A long-term case history study.

Western Society of Naturalists Annual Meeting, Monterey, USA.
Lecture: Sexual reproduction in reef corals: Red Sea versus the Great Barrier Reef.
Fifth International Coral Reef Congress, Tahiti.
Invited lecture: Seasonal changes in growth rate of a Red Sea coral population.
Second Conference of the International Society of Reef Studies, Nice, France.
Plenary lecture: Reproductive patterns of Red Sea corals.

International Helgoland Symposium: “Diseases of Marine Organisms”.

Biologisch Anstalt Helgoland, Helgoland, Germany F.R.
Invited lecture: Tumor formations in scleractinian corals.

Great Barrier Reef Conference. James Cook University and the
Australian Institute of Marine Science (AIMS), Townsville, Australia.
Plenary lecture: Community structure and life history strategies of scleractinian corals.

Workshop on Evolution and Maintenance of Modern Coral Distributions.
Australian Institute of Marine Science, Townsville, Australia.
Invited lecture: Reproductive cycle of Stylophora pistillata in the Great Barrier

Reef and the Red Sea.


International Conference on Marine Science in the Red Sea. Al-Ghardaqa

Marine Biological Station, Egypt.
Lecture: Seasonal changes in growth rates of a Red Sea coral population.


U.S. National Academy of Sciences - Update workshop on: “Oil in the sea: Inputs, Fates and Effects”.
Invited referee in the section of Effects of Petroleum Hydrocarbons on Coral Reefs. Clearwater, Florida, USA.

Fourth International Coral Reef Symposium, Manila, Philippines.
Lecture: Life history strategies of boring bivalves in corals

Second International Symposium on Biology and Management of Mangroves and Tropical Shallow Water Communities. The Western Society of Naturalists and the University of Papua New Guinea. Port Moresby, Madang, Papua New Guinea.
Plenary lecture: Competition for space among coral reef sessile organisms.


UNESCO workshop on Marine and Coastal Processes in the Pacific: Ecological aspects of coastal zone management;
Mutupore Island Research Centre, Port Moresby, Papua New Guinea
Panel leader: Degradation of the environment.

Australian Institute of Marine Science (AIMS) Workshop on Coral Reefs;
Cape Ferguson, Townsville, Australia
Lecture: Life history strategy of a Red Sea coral population.
Second International Congress of Ecology, Jerusalem.

Tenth International Congress of Sedimentology, Jerusalem.
Field leader: “Pleistocene and Recent coral reefs and coastal sedimentation in
the Gulf of Eilat”.


Third International Coral Reef Symposium, Univ. of Miami, Florida.
   1. Seasonal occurrence of benthic-algae communities and grazing regulation by

       seaurchins at the coral reefs of Eilat, Red Sea.
   2. Harmful effects of chronic oil pollution on a Red Sea scleractinian coral population.


International Helgoland Symposium: Ecosystem Research. Biologische Anstalt Helgoland - Helgoland, Germany F.R.
Invited lecture: Quantitative characteristics of community structure of stony corals,
soft corals and algae in the northern Gulf of Eilat, Red Sea.

Third International Symposium on Coelenterate Biology, Victoria, B.C., Canada.
Lecture: Settlement, mortality and recruitment in a Red Sea scleractinian coral population.

Second International Symposium on Coral Reefs, The Great Barrier Reef, Australia.
Plenary lecture: On the possible use of plotless methods for quantitative studies of
benthic communities of coral reefs.

The Penrose Conference on Marine Ecology and Paleo-ecology, Monterey, California.
Lecture: Community structure and species diversity of hermatypic corals.

Symposium of Regional Variation of Indian Ocean Coral Reefs. The Royal Society and Zoological Society of London, London.
Invited lecture: The coral reefs of Eilat, Red Sea.

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